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Pine Needles is another buck that escaped the Sandleford Warren when men sprayed poison gas into the holes. Pine Needles escaped into the forest with Ash, Butterbur, and Nose-in-the-Air. He was one of eight that escaped in total, with the others being Toadflax, Bluebell, Pimpernel, and Captain Holly.

Rabbits
Cross-media
Sandleford Warren
BigwigBlackberryBluebellCaptain HollyDandelionFiverHawkbitHazelHazel and Fiver's MotherPipkinPimpernelSilverThrearahToadflax

Nuthanger Farm
BoxwoodCloverHaystackLaurel
Cowslip's Warren
CowslipKingcupLaburnumSilverweedStrawberry
Efrafa
BlackavarCaptain CampionCaptain ChervilCaptain OrchisCharlockGeneral WoundwortHyzenthlayRagwortSainfoinThethuthinnangThistleVervainVilthuril
Folklore
Black Rabbit of InléEl-ahrairahHufsaRabscuttle

Books only
Introduced in Watership Down
AcornAshAvensBartsiaBetonyBuckthornButterburCaptain BuglossCaptain MallowColtsfootFlescaGroundselMarjoramMoneywortNelthiltaNightshadeNildro-hainNose-in-the-AirPine NeedlesScabiousSnowdropSpeedwellThrayonlosaThrearThunder

Introduced in Tales from Watership Down
BurdockCelandineCrowfootCrowlaFescueFlyairthFoxgloveHeartseaseKing Fur-RociousKnapweedLemistaLoosestrifeMelsaMianMilmownNyreemPeertonPrakeQuiensRithlaSandwortStitchwortStonecropThrennionTindraWoodruff

AspenBlueskyButtercupCaptain BroomFelsparFlaxGentianGiliaGorseGraniteHeatherHickoryLarchMallowMarigoldMossPrimroseRaincloudRedShaleSnowdropSpartinaSpeaker of the PastLaurel
Film/Miniseries only
Dewdrop (Miniseries) • Nettle (Miniseries) • Violet (film)
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